I just saw your other thread about this problem and you mentioned that you are using Tap Water (wish you had put all the information in the same thread as I now have more information to add).

I got this problem when I ran out of PH tested pre-mixed food/water and had to feed my ladies straight tap water for 4 days. I thought the "lipped" leaves and brown spots were related to the same problem, but after reading your post I now think the problems are caused by seperate issues.

I went back to my normal feeding schedule and overhauled my ventilation system at the same time. This worked for me.